(19)

(11)

EP 0 426 840 A1

(12)

(43) Date of publication: 15.05.1991 Bulletin 1991/20 (21) Application number: 90909228.0 (22) Date of filing: 17.05.1990

(51) International Patent Classification (IPC):  C12N 15/ 09( . ) C12N 15/ 866( . ) C12N 15/ 68( . ) (86) International application number: PCT/US1990/002814 (87) International publication number: WO 1990/014428 (29.11.1990 Gazette 1990/27)

(84) Designated Contracting States: AT BE CH DE DK ES FR GB IT LI LU NL SE (30) Priority: 17.05.1989 US 19890353847 (71) Applicant: THE UNIVERSITY OF GEORGIA RESEARCH FOUNDATION, INC. Athens, Georgia 30602 (US)	(72) Inventor: MILLER, Lois, K. Athens, GA 30605 (US) (74) Representative: Fisher, Adrian John  CARPMAELS & RANSFORD 43 Bloomsbury Square London WC1A 2RA London WC1A 2RA (GB)