(19)

(11)

EP 0 819 174 A1

(12)

(43) Date of publication: 21.01.1998 Bulletin 1998/04 (21) Application number: 96909256.0 (22) Date of filing: 09.04.1996

(51) International Patent Classification (IPC):  A01H 1/ 00( . ) C07K 14/ 37( . ) C12N 1/ 21( . ) C12N 15/ 09( . ) C12N 15/ 82( . ) C12R 1/ 01( . ) A01H 5/ 00( . ) C07K 14/ 415( . ) C12N 5/ 10( . ) C12N 15/ 29( . ) C12Q 1/ 68( . ) (86) International application number: PCT/GB1996/000849 (87) International publication number: WO 1996/031608 (10.10.1996 Gazette 1996/45)

(84) Designated Contracting States: AT BE CH DE DK ES FI FR GB GR IE IT LI NL PT SE (30) Priority: 07.04.1995 GB 19950007232 (71) Applicant: JOHN INNES CENTRE INNOVATIONS LIMITED Norwich, Norfolk NR4 7UH (GB)

(72) Inventors: JONES, Jonathan Dallas George Norwich, Norfolk NR4 6SG (GB) PARKER, Jane Norwich, Norfolk NR 2 3RW (GB) COLEMAN, Mark Norwich, Norfolk NR2 3BN (GB) DANIELS, Michael John Norwich, Norfolk NR4 7QY (GB) SZABO, Véronique, University of California Davis, CA 95616 (US) (74) Representative: Walton, Seán Malcolm, et al  MEWBURN ELLIS, York House, 23 Kingsway London WC2B 6HP London WC2B 6HP (GB)