(19)

(11)

EP 0 858 511 A1

(12)

(43) Date of publication: 19.08.1998 Bulletin 1998/34 (21) Application number: 96937322.0 (22) Date of filing: 31.10.1996

(51) International Patent Classification (IPC):  A01H 5/ 00( . ) C12N 5/ 10( . ) C12N 15/ 09( . ) C12N 15/ 82( . ) A01H 5/ 10( . ) C12N 9/ 10( . ) C12N 15/ 54( . ) (86) International application number: PCT/EP1996/004747 (87) International publication number: WO 1997/016559 (09.05.1997 Gazette 1997/20)

(84) Designated Contracting States: AT BE CH DE DK ES FI FR GB GR IE IT LI LU MC NL PT SE (30) Priority: 31.10.1995 EP 19950402425 (71) Applicants: PLANT GENETIC SYSTEMS N.V. 9000 Gent (BE) NATIONAL RESEARCH COUNCIL CANADA Ottawa Ontario K1A OR6 (CA)

(72) Inventors: VAN AUDENHOVE, Katrien B-8310 Assebroek (BE) MARILLIA, Elizabeth Saskatoon, Saskatchewan S7K 3J1 (CA) PEFEROEN, Marnix B-9850 Nevele (BE) GROOTWASSINK, Jan, Willem, Derk Saskatoon, Saskatchewan S7H 3A6 (CA) REED, Darwin, Wilfred Saskatoon, Saskatchewan S7M 4J9 (CA) HEMMINGSEN, Sean, Mathias Saskatoon, Saskatchewan S7H 3A2 (CA) KOLENOVSKY, Allan, Dale Saskatoon, Saskatchewan S7K 5C4 (CA) UNDERHILL, Edward, Wesley Victoria, British Columbia V8V 2T4 (CA) MACPHERSON, James, Malcolm Saskatoon, Saskatchewan S7L 3V8 (CA) (74) Representative: Desaix, Anne, et al  Ernest Gutmann - Yves Plasseraud S.A. 3, rue Chauveau-Lagarde 75008 Paris 75008 Paris (FR)