(19)

(11)

EP 1 135 507 A1

(12)

(43) Date of publication: 26.09.2001 Bulletin 2001/39 (21) Application number: 99958511.0 (22) Date of filing: 02.12.1999

(51) International Patent Classification (IPC)7: C12N 15/53, C12N 15/54, C12N 15/55, C12N 15/60, C12N 15/11, C12N 15/82, C12N 9/10, C12N 9/18, C12N 9/88, C12N 9/04, C12P 7/62, C12Q 1/68, C07K 16/40, A01H 5/00, G01N 33/50 (86) International application number: PCT/NL9900/737 (87) International publication number: WO 0032/789 (08.06.2000 Gazette 2000/23)

(84) Designated Contracting States: AT BE CH CY DE DK ES FI FR GB GR IE IT LI LU MC NL PT SE Designated Extension States: AL LT LV MK RO SI (30) Priority: 02.12.1998 EP 98204018 12.03.1999 EP 99200739 (71) Applicant: CENTRUM VOOR PLANTENVEREDELINGS- EN REPRODUKTIEONDERZOEK 6708 PB Wageningen (NL)	(72) Inventors: AHARONI, Asaph Tel-Aviv (IL) LÜCKER, Joost 5991 DA Baarlo (NL) VERHOEVEN, Harrie, Adrianus NL-5694 LB Breugel (NL) VAN TUNEN, Arjen, Johannes NL-6708 NA Wageningen (NL) O'CONNELL, Ann, Patricia NL-2633 JH Nootdorp (NL) (74) Representative: Prins, Adrianus Willem et al Vereenigde,Nieuwe Parklaan 97 2587 BN Den Haag 2587 BN Den Haag (NL)