(19)

(11)

EP 1 266 017 A1

(12)

(43) Date of publication: 18.12.2002 Bulletin 2002/51 (21) Application number: 00926838.4 (22) Date of filing: 23.03.2000

(51) International Patent Classification (IPC)7: C12N 15/74 // C12R1:225 (86) International application number: PCT/EP0003/099 (87) International publication number: WO 0107/1011 (27.09.2001 Gazette 2001/39)

(84) Designated Contracting States: AT BE CH CY DE DK ES FI FR GB GR IE IT LI LU MC NL PT SE Designated Extension States: AL LT LV MK RO SI (71) Applicant: INSTITUT NATIONAL DE LA RECHERCHE AGRONOMIQUE (INRA) 75338 Paris Cédex 07 (FR)	(72) Inventors: GORY, Laure F-43700 Coubon (FR) MONTEL, Marie-Christine F-15000 Aurillac (FR) ZAGOREC, Monique F-78350 Jouy En Josas (FR) (74) Representative: Vialle-Presles, Marie José et al Cabinet Orès,6, Avenue de Messine 75008 Paris 75008 Paris (FR)