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(11)

EP 3 013 939 A1

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(43) Date of publication: 04.05.2016 Bulletin 2016/18 (21) Application number: 14744471.5 (22) Date of filing: 25.06.2014

(51) International Patent Classification (IPC):  C12N 1/13(2006.01) C12N 9/02(2006.01) C12N 9/12(2006.01) C12P 7/64(2006.01) C12R 1/89(2006.01) C12N 9/04(2006.01) C12N 9/10(2006.01) C12N 9/16(2006.01) C12N 15/79(2006.01) (86) International application number: PCT/EP2014/063393 (87) International publication number: WO 2014/207043 (31.12.2014 Gazette 2014/53)

(84) Designated Contracting States: AL AT BE BG CH CY CZ DE DK EE ES FI FR GB GR HR HU IE IS IT LI LT LU LV MC MK MT NL NO PL PT RO RS SE SI SK SM TR Designated Extension States: BA ME (30) Priority: 25.06.2013 DK 201370354 (71) Applicant: Cellectis 75013 Paris (FR)	(72) Inventors: DUCHATEAU, Philippe F-91210 Draveil (FR) DABOUSSI, Fayza F-77500 Chelles (FR) SOURDIVE, David F-92300 Levallois-Perret (FR) EPINAT, Jean-Charles F-93260 Les Lilas (FR) (74) Representative: Zacco Denmark A/S  Arne Jacobsens Allé 15 2300 Copenhagen S 2300 Copenhagen S (DK)