(19)

(11)

EP 3 468 589 A1

(12)

(43) Date of publication: 17.04.2019 Bulletin 2019/16 (21) Application number: 17730747.7 (22) Date of filing: 09.06.2017

(51) International Patent Classification (IPC):  A61K 39/12(2006.01) C12N 15/86(2006.01) (86) International application number: PCT/EP2017/064183 (87) International publication number: WO 2017/212048 (14.12.2017 Gazette 2017/50)

(84) Designated Contracting States: AL AT BE BG CH CY CZ DE DK EE ES FI FR GB GR HR HU IE IS IT LI LT LU LV MC MK MT NL NO PL PT RO RS SE SI SK SM TR Designated Extension States: BA ME Designated Validation States: MA MD (30) Priority: 10.06.2016 WO PCT/EP2016/305697 (71) Applicant: Ceva Santé Animale 33500 Libourne (FR)	(72) Inventors: SATO, Takanori Fujisawa-shi Kanagawa 221-0012 (JP) SAITOH, Shuji Yokohama 235-0045 (JP) KOMIYA, Yasutoshi Yokohama 230-0012 (JP) (74) Representative: Cabinet Becker et Associés  25, rue Louis le Grand 75002 Paris 75002 Paris (FR)