(19)

(11)

EP 4 077 652 A1

(12)

(43) Date of publication: 26.10.2022 Bulletin 2022/43 (21) Application number: 20842218.8 (22) Date of filing: 18.12.2020

(51) International Patent Classification (IPC):  C12N 9/22(2006.01) C12N 15/10(2006.01) C12N 9/78(2006.01) (52) Cooperative Patent Classification (CPC): C12N 9/22; C12N 9/78; C12N 15/102; C12N 2310/20; C12N 15/8213; C12N 9/93; C12N 15/8274; C12N 9/1085; C12N 15/8212 (86) International application number: PCT/EP2020/087344 (87) International publication number: WO 2021/123397 (24.06.2021 Gazette 2021/25)

(84) Designated Contracting States: AL AT BE BG CH CY CZ DE DK EE ES FI FR GB GR HR HU IE IS IT LI LT LU LV MC MK MT NL NO PL PT RO RS SE SI SK SM TR Designated Extension States: BA ME Designated Validation States: KH MA MD TN (30) Priority: 20.12.2019 EP 19306747 (71) Applicant: Limagrain Europe 63360 Saint-Beauzire (FR)	(72) Inventors: PAUL, Wyatt 63720 Chappes (FR) NOGUÉ, Fabien 78026 Versailles (FR) GUYON, Anouchka 78026 Versailles (FR) PERROUD, Pierre-François 78026 Versailles (FR) (74) Representative: Flesselles, Bruno F.G.  BF IP 36 rue Jean de la Fontaine 75016 Paris 75016 Paris (FR)